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Periodicity of strong nucleosome positioning sites around the chicken adult β-globin gene may encode regularly spaced chromatin
Journal article   Peer reviewed

Periodicity of strong nucleosome positioning sites around the chicken adult β-globin gene may encode regularly spaced chromatin

C Davey, S Pennings, G Meersseman, Tim J Wess and James Allan
Proceedings of the National Academy of Sciences of the United States of America, Vol.92(24), pp.11210-11214
1995
url
https://doi.org/10.1073/pnas.92.24.11210View
Published Version

Abstract

Positioned nucleosomes contribute to both the structure and the function of the chromatin fiber and can play a decisive role in controlling gene expression. We have mapped, at high resolution, the translational positions adopted by limiting amounts of core histone octamers reconstituted onto 4.4 kb of DNA comprising the entire chicken adult β-globin gene, its enhancer, and flanking sequences. The octamer displays extensive variation in its affinity for different positioning sites, the range exhibited being about 2 orders of magnitude greater than that of the initial binding of the octamer. Strong positioning sites are located 5' and 3' of the globin gene and in the second intron but are absent from the coding regions. These sites exhibit a periodicity (≃200 bp) similar to the average spacing of nucleosomes on the inactive β-globin gene in vivo, which could indicate their involvement in packaging the gene into higher-order chromatin structure. Overlapping, alternative octamer positioning sites commonly exhibit spacings of 20 and 40 bp, but not of 10 bp. These short-range periodicities could reflect features of the core particle structure contributing to the pronounced sequence- dependent manner in which the core histone octamer interacts with DNA.

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