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Evolution of higher torque in Campylobacter-type bacterial fagellar motors
Journal article   Open access   Peer reviewed

Evolution of higher torque in Campylobacter-type bacterial fagellar motors

Bonnie L Chaban, Izaak Coleman and Morgan Beeby
Scientific Reports, Vol.8, 97
2018
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https://doi.org/10.1038/s41598-017-18115-1View
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Abstract

Understanding the evolution of molecular machines underpins our understanding of the development of life on earth. A well-studied case are bacterial fagellar motors that spin helical propellers for bacterial motility. Diverse motors produce diferent torques, but how this diversity evolved remains unknown. To gain insights into evolution of the high-torque ε-proteobacterial motor exemplifed by the Campylobacter jejuni motor, we inferred ancestral states by combining phylogenetics, electron cryotomography, and motility assays to characterize motors from Wolinella succinogenes, Arcobacter butzleri and Bdellovibrio bacteriovorus. Observation of ~12 stator complexes in many proteobacteria, yet ~17 in ε-proteobacteria suggest a "quantum leap" evolutionary event. Campylobacter-type motors have high stator occupancy in wider rings of additional stator complexes that are scafolded by large proteinaceous periplasmic rings. We propose a model for motor evolution wherein independent innerand outer-membrane structures fused to form a scafold for additional stator complexes. Signifcantly, inner- and outer-membrane associated structures have evolved independently multiple times, suggesting that evolution of such structures is facile and poised the ε-proteobacteria to fuse them to form the high-torque Campylobacter-type motor

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